In good old RuneQuest it was a kind of spiritual thing - through adventuring (questing) you strove to improve yourself, in skill and holiness, in order to emulate your god and thus attune yourself with the Runes (primal natural principles) they embodied.
Mongoose failed to see that, and insensitively turned Runes into just another magic treasure to be looted off the dead bodies of the monsters/victims, D&D-style. Pah.
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